Megalosaurus is also unusual in that it has broad pubes

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Megalosaurus is also unusual in that it has broad pubes, like early dinosaurs. (Walking dinosaur costume)This once led me to believe that it was the most archaic of theropods,e and Peter Galton and James ("Dinosaur") Jensen to conclude that megalosaurs and some other big theropods evolved completely separately from small theropods. But megalosaurs share too many characters with other theropods for this to be true. I now believe the broad hips are a secondary reversal to the old condition. Eustreptospondylians and metriacanthosaurs are a bit more advanced than megalosaurs in the above listed characters, and less abenant. Theropods like these may have been the ancestors of more advanced theropods. Metriocanthoscurus is especially important in helping us ascertain the evolutionary sequence, because this well-preserved form has a large, expanded pubic tip, or boot, and the shorter anterior trunk ribs that go with the start of an avian lung system.

 

We now come to avetheropods.to These are very birdlike dinosaurs, even for theropods, having many avian adaptations. For example, the lower jaw is hinged at its midlength, and many birds retain this in a modified form. At the back of the roof of the mouth the pterygoid bone has a reduced connection between its two main sections; this would become a hinge in many birds (Realistic dinosaur costume). The front trunk ribs are short and slender, and the back ones are long in order to operate abdominal air sacs (see Figure 4-8, page 104). The shoulder blade is very narrow, also a birdlike feature, as are the big inner wrist bone and three-fingered hand (Animatronic dinosaur). The pubic boot is very large, and the outer process of the femoral head is very tall. Figure 8-8 shows that the ankle has another very avian adaptation, a tall ascending process of the astragalus.' Finally, the upper part of the central cannon bone is narrower tha4 in paleotheropods (Figure 8-9). The progressive narrowing of the central cannon bone in theropods is sometimes portrayed as either strongly compressed at its upper end, or not compressed at all. Actually it is the degree of narrowing that is very important.

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